people / Ray, Chris
Ray, Chris


Chris Ray

Office: 303-489-8863



Chris Ray studies and models the dynamics of plant and animal populations, focusing especially on threatened species with fragmented populations. Recent projects include hierarchical modeling of point count data, analyzing the role of metacommunity dynamics in plant community assembly, and modeling the spatial and temporal dynamics of disease in mammal communities. Her long-term project involves research on the American pika, aimed at understanding climatic influences on the distribution of this species throughout western North America.

Disciplines: Animal ecology




Chris Ray Biography: INSTAAR

Chris Ray Biography: IBP

Outreach: Mammal Class, Rocky Mountain National Park

Front Range Pika Project

Chris Ray's CV


Identification of a contact zone and hybridization for two subspecies of the American pika (Ochotona princeps) within a single protected area.

Castillo Vardaro, J.A., Epps, C.W., Frable, B.W., Ray, C. , PLoS ONE, 13, 7, DOI: 10.1371/journal.pone.0199032,


Individual-based analysis of hair corticosterone reveals factors influencing chronic stress in the American pika.

Waterhouse, M.D., Sjodin, B., Ray, C., Erb, L., Wilkening, J., Russello, M.A., Ecology and Evolution, 1-10, DOI: 10.1002/ece3.3009,


Rodent–pika parasite spillover in western North America.

Foley, P. , Roth, T., Foley, J., Ray, C., Journal of Medical Entomology,, 1-7, DOI: 10.1093/jme/tjx085,


Characterizing predictors of survival in the American pika (Ochotona princeps).

Wilkening, JL, Ray, C, Journal of Mammalogy, 1-10, DOI: 10.1093/jmammal/gyw097,


When can we measure stress noninvasively? Postdeposition effects on a fecal stress metric confound a multiregional assessment.

Wilkening, JL, Ray, C, Varner, J, Ecology and Evolution, 6, 2, 502-513, DOI: 10.1002/ece3.1857,


Of plants and pikas: evidence for a climate-mediated decline in forage and cache quality.

Bhattacharyya, S., Ray, Chris, Plant Ecology & Diversity, 8, 5-6, 781-794, DOI: 10.1080/17550874.2015.1121520,


Alpine biodiversity and assisted migration: the case of the American pika (Ochotona princeps).

Wilkening, JL, Ray, C, Ramsay. N, Klingler, K, Biodiversity, 16, 4, 224-236, DOI: 10.1080/14888386.2015.1112304,


Parks, pikas, and physiological stress: Implications for long-term monitoring of an NPS climate-sensitive sentinel species.

Wilkening, JL, Ray, C, Park Science, 32, 1, 42-48,


Relating sub-surface ice features to physiological stress in a climate sensitive mammal, the American pika (Ochotona princeps).

Wilkening, Jennifer L., Ray, Chris, Varner, Johanna, PLOS one, 10, 3, e0119327, DOI: 10.1371/journal.pone.0119327,


Children’s book series and associated curricula support elementary education and outreach in water resources.

Ray, C., McKnight, D. M., Bidwell, M. D., Fourment, T., Flanagan Pritz, C., Rinehart, A. H., Plant Ecology & Diversity, DOI: 10.1080/17550874.2015.1050711,


Determinants of pika population density versus occupancy in the Southern Rocky Mountains.

Erb, L.P., Ray, C., Guralnick, R., Ecological Applications, 24, 3, 429-435, DOI: 10.1890/13-1072.1,


Stress hormone concentration in Rocky Mountain populations of the American pika (Ochotona princeps).

Wilkening, J.L., Ray, C., Sweazea, K.L., Conservation Physiology, DOI: 10.1093/conphys/cot027,


Retreat of the American pika: Up the mountain or into the void?.

Ray, C., Beever, E., Loarie, S., Wildlife Conservation in a Changing Climate, Chicago, University of Chicago Press, Brodie, J.F., Post, E., and Doak, D.F., 245-270,

CU Boulder

This material is based upon work supported by the National Science Foundation under Cooperative Agreement #DEB-1637686. Any opinions, findings, conclusions, or recommendations expressed in the material are those of the author(s) and do not necesarily reflect the views of the National Science Foundation.

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